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Suc is a disaccharide that can be hydrolyzed into Glc and Fru by specific Inv enzymes found in three distinct sub-cellular localizations: the cell wall (CW-Inv), the cytoplasm (C-Inv), or the vacuole (V-Inv) 77 .The fate of Suc as a carbon source during natural leaf senescence is the export to the sink organs via the apoplast 16,74 or the import into the vacuole through the tonoplast, where V-Inv activities can catalyze its hydrolysis

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